Thank you for your question. Something you are coding in the .htaccess is causing the 500 error, here is a link to our 500 error troubleshooting guide. Essentially, the server returns a 500 error when there is no other error to display, you can however create a custom error page.
Wordpress has a debugging mode, which will allow it to enable and display errors.
I recommend leaving the .htaccess as is, since Wordpress requires it to function.
You cannot add the PHP rules to the .htaccess, since this will not work on a suPHP server, which is what we run.
You can turn these PHP error reports on in your PHP.ini file, then make it recursive.
If you have any further questions, feel free to post them below.
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